The study of all the available skulls allows us to review the systematic relationships of the longirostrine Steneosaurus from the Middle Jurassic of western Europe. Up to now, Aalenian and Bajocian deposits have not yielded any significant Steneosaurus remain. In the Bathonian, the only valid longirostrine species, S. megistorhynchus, is known in the Britain-Normandy Basin, the Poitou and the Lorraine. In the Callovian, most of the longirostrine Steneosaurus remains can be attributed to the species S. leedsi. Nevertheless, some remains from the Middle Callovian of Poitou (France) show important differences with S. leedsi. A new Steneosaurus species, only known in Poitou, is created and named S. pictaviensis. The specific characters are carried by the skull (preorbital pit well marked, orbit and ptetygoid fossae shapes), by the mandible (symphysis shape) and by the teeth (ornamentation). S. megistorhynchus is probably situated near the stem of the Callovian species but remains from the Bathonian and Lower Callovian are very scarce and it is very difficult to precise the phylogenetic relationships between the longirostrine species of the Middle Jurassic.
  

An increasing fossil record of dinosaur eggs and eggshells allows putting ootaxa within a chronostratigraphic framework, in order to study their distribution pattern leading eventually to their use as biochronological markers. For these purposes, high-quality data exists in four major regions; North America, South America, Europe and Asia (Central Asia and India). Most of the highly diverse dinosaur egg record has been dated as Latest Cretaceous in age (Campanian-Maastrichtian), reaching the Cretaceous-Tertiary boundary closer than the dinosaur bone record. However, dating continental sections is problematic and need to be carefully verified, as it appears when comparing the European dinosaur eggshell record from two well-studied areas. Ootaxa distribution in both sides of the Pyrenees (Tremp and Aix basins) shows comparable oospecies successions, but different chronology. This disagreement probably indicates that one or both successions have a wrong chronostratigraphic calibration.  

The genus Cryptomeryx SCHLOSSER, 1886, inusited during a long period, has been discovered in Lower and Middle Oligocene localities of the Quercy region (South-West France). This new material, as well as specimens from the old collections referred to Cryptomeryx, are described; their study, allows us precising the definition of the genus, and confirming its allocation to the Tragulidae family. The type species of the genus, Crypmmeryx gaudryi (= Lophiomeryx gaudryi FILHOL, 1877), occurs in several localities at the base of the Middle Oligocene (Itardies, La Plante 2, Roqueprune 2, Soulce, Herrlingen 1). The new species C. matsoui n. sp. has been defined in the older locality of Mas de Got (top of Lower Oligocene). It is possible that the species Pseudamphimeryx decedens STEHLIN, 1910 pertains to the same genus. Also to the Tragulids must be referred the monospecific genus Iberomeryx (I. parvus GABOUNIA, 1964) from Upper Oligocene of Benara (Georgie, URSS), with which Cryptomeryx is related. These genera are not direct ancestors of Miocene tragulids; their occurrence in the Western European Oligocene results from a first immigration wawe of the family. These Tragulids are one of the most archaic groups of Ruminants. They are probably derived from a primitive stock which had acquired in Asia the selenodont condition of the dentition. 

A new Devonian vertebrste locality has been discovered in 1976 in the basal part of the Khush-Yeilagh Formation in the eastern Alborz Mountains of Iran. The fossils occur in a band one centimeter thick which is identifiable at other outcrops in the area. A preliminary study of the remains has yielded the following faunal list : Placodermata (Phlyctaeniida indet.. Groenlandaspididae indet.. Coccosteidae indet., Holonema sp., Ptyctodontida indet., Antiarcha indet., Bothriolepis cf. kwangtungensis), Elasmobranchii (Ctenacanthidae indet.), Acanthodii (Gyracanthus sp., « Onchus » overathensis, Ischnacanthiforme indet.), Dipnoi (? Dipteridae indet.), « Crossopterygii ›› (Onychodus cf. sigmoides, Holoptychiidae indet., Osteolepididae indet.). This fauna is older than the Middle Givetian and probably of Middle or Lower Eifelian age. Owing to its diversity, it may prove possible to use it in the future as a reference fauna for the study of the Devonian vertebrates in Central Asia and the Middle East. 

Based on fossil discoveries and phylogenetic studies, an Eocene Asian origin for hystricognathous rodents and anthropoid primates has gained strong support in recent years. The two groups then invaded both Africa and South America, which promoted their evolutionary success. However, the fossil record has so far failed to constrain the nature and precise timing of these pivotal dispersal events. In Africa, given the apparent absence of hystricognaths and anthropoids in early to early middle Eocene localities, it is suggested that these mammal groups dispersed from Asia to Africa sometime during the middle Eocene. In this paper, we report the discovery of several isolated teeth of a rodent from a new vertebrate locality situated in central Tunisia (Djebel el Kébar, KEB-1), dating from the late middle Eocene (Bartonian, ~39.5 Myr). These fossils document a diminutive new species of Protophiomys (P. tunisiensis nov. sp.), a basal genus of hystricognathous rodents which is well known from several North African mammalian-bearing localities of the end of the Eocene. The teeth of P. tunisiensis display a suite of anatomical details comparable with those observed in the other species of the genus, but with a lesser degree of development. Such an apparent primitive evolutionary stage is corroborated by the greater antiquity of this Tunisian species. P. tunisiensis nov. sp. is so far the most ancient representative of hystricognaths in Africa. However, it can be expected that hystricognaths were already present on that landmass given the new data on early caviomorphs recently reported from South America (at ~41 Myr). The arrival of hystricognaths in Africa from South Asia certainly predates the depositional period of the Kébar sediments, but perhaps not by much time. 

Eosimiid and amphipithecid primates document a long and significant history of primate evolution throughout the Eocene in Southeast Asia. Despite the absence of a comprehensive post-Eocene fossil record, it was generally hypothesized that both families left no descendant in Asia. Recently, two new small-bodied taxa, Bugtipithecus and Phileosimias, have been recovered in early Oligocene coastal deposits from the Bugti Hills (Balochistan, central Pakistan) and referred to the families Amphipithecidae and Eosimiidae, respectively, on the basis of dental fossil remains. In this paper, we provide more exhaustive description, comparison, and discussion of these taxa. As for tarsiid and sivaladapid primates, the persistence of eosimiids and amphipithecids into the Oligocene clearly demonstrates that low latitudes of South Asia provided a continuous access to tropical refugia during the climatic deterioration characterizing the late Eocene-early Oligocene interval, which was seemingly lethal for primate communities elsewhere across the Holarctic continents. As a contribution to the ongoing phylogenetic debates regarding the position of eosimiids and amphipithecids on the primate family tree, we have performed a cladistic analysis in a high-level primate systematic context in order to assess the position and the role of these new taxa in that phylogenetic issue. Our results support the view according to which eosimiids and amphipithecids (and by extension Phileosimias and Bugtipithecus, respectively) are stem anthropoids. These fossils from Pakistan document an unsuspected Oligocene phase of the evolutionary history of anthropoid primates in southern Asia, which clearly enhances the extent of the anthropoid radiation in this province during the Paleogene. Several phylogenetic and paleobiogeographic aspects are discussed, notably the intra- and inter-relationships between Paleogene Asian and Afro-Arabian anthropoids, and the resulting potential dispersal models between both land-masses during the Paleogene. 

Anthracotheres from the Pondaung Formation, Myanmar, are considered as one of the most primitive artiodactyl groups and they represent the oldest known record in the world. Thus, the understanding of this group has numerous implications for evolutionary biology and biochronological correlations. However, the systematlcs of these mammals has been interpreted in different ways, and the main debate focuses on the number of taxa represented in the Pondaung Formation. The revised taxonomy proposed here is mainly based on the relative development of the upper molar W-shaped ectoloph, system of crests and stylar cusps, and on body size. On the basis of these characters, they are classified into four genera including six different species. Two well-known genera, Anthracotherium and Anthracokeryx, are validated and more precisely diagnosed. Anthracokeryx possesses a better developed W-shaped ectoloph, system of crests and stylar cusps than Anthracotherium, which displays notable differences with the more derived representatives of this genus. Both of these Pondaung genera show evidence for sexual dimorphism. However, the incompleteness of fossil material fueled a debate concerning the status of two additional Pondaung anthracotheres, Siamotherium and Anthracohyus. The latter genus is of uncertain affinities, but it has been considered as a hippopotamid ancestor. Despite new material attributed to these two forms, additional discoveries are still required to establish their taxonomic status. The hypothesis that Southeast Asia was the centre of origin of Anthracotheriidae is supported by the retention of numerous primitive dental characters in these taxa and by the antiquity of the Pondaung Formation, to which an age of 37 My is now generally accepted. 

An isolated right costal 1 from the Late Cretaceous Massecaps locality (Cruzy, Hérault, southern France) is assigned to Dortoka vasconica (Dortokidae). This find adds a new element to the Late Cretaceous turtle fauna of Cruzy and further supports the hypothesis that two distinct lineages of Dortokidae were present in Europe during the Late Cretaceous-Paleogene due to geographical isolation.
  

Micro-CT imaging is an increasingly popular method in paleontology giving access to internal structures with a high resolution and without destroying precious specimens. However, its potential for the study of hypoplasia defects has only recently been investigated. Here, we propose a preliminary study to test whether hypoplastic defects can be detected with micro-CT (μCT) scan and we assess the costs and benefits of using this method instead of naked eye. To do so, we studied 13 fossil rhinocerotid teeth bearing hypoplasia from Béon 1 (late early Miocene, Southwestern France) as positive control and 11 teeth of the amynodontid Cadurcotherium (Oligocene, Phosphorites du Quercy, Southwestern France), for which enamel was partly or totally obscured by cement. We showed that all macroscopically-spotted defects were retrieved on 3D reconstructions and selected virtual slices. We also detected additional defects using μCT scan compared to naked eye identification. The number of defects detected using μCT was greater in the Cadurcotherium dataset (paired-sample Wilcoxon test, p-value = 0.02724) but not for our control sample (paired-sample Wilcoxon test, p-value = 0.1171). Moreover, it allowed for measuring width and depth of the defects on virtual slices (sometimes linked to stress duration and severity, respectively), which we could not do macroscopically. As μCT imaging is both expensive and time consuming while not drastically improving the results, we recommend a moderate and thoughtful use of this method for hypoplasia investigations, restricted for instance to teeth for which enamel surface is obscured (presence of cement, uncomplete preparation, or unerupted germs). 

The Late Miocene site of Gritsev (MN 9, Ukraine) has yielded a very rich bat fauna, the remains of which are well preserved. Compared to other Neogene bat assemblages of Europe, the Gritsev bat community is unique in preserving plecotine bats, which are rare from Neogene sites. Some peculiar and new bat species, including a large plecotin Otonycteris, already were described from the Gritsev mammal site. Here we report new records of small plecotin bats from Gritsev, including a new taxon, Barbastella maxima nov. sp. This is the earliest reliable fossil record of this genus and it differs from more recent species of Barbastella in being considerably larger. The evolutionary patterns in the odontology within the tribe Plecotini, supported by biostratigraphical distribution of fossil records of Plecotus are discussed. The morphological peculiarities of the new fossils of plecotine bats from Gritsev are discussed in connection with its possible taxonomical affinity. 

Three new fossiliferous localities, two of karstic origin, Castelnou 3 and Font Estramar, respectively Late Upper Miocene and Lower Pliocene, and one of lacustrine origin, Thuir, Lower Pliocene, add data about the transition between Miocene and Pliocene faunas of rodents in southern France. An unexpected association of taxa was present in the late Upper Miocene, including between others, Myocricetodon, Hispanomys, Ruscinomys, Cricetus barrierei, Promimomys and a new species of Stephanomys, S. dubari nov. sp. Myocricetodon is still known in the Lower Pliocene. It is shown that the large field-mice known since the Late Upper Miocene belong to two different lineages, on one side, A. jeanteti, on the other side, A. gudrunae followed by A. gorafensis. Biochronological and biogeographical implications are discussed. 

An assemblage of micromammals is reported from California Wash, a fossil bearing continental deposit in the San Pedro Valley, Arizona, late Blancan in age. Cricetid and Arvicolid rodents are richly represented, including four and two species, respectively. This study mainly focuses on Sígmodon, the most abundant form. The sample of Sigmodon is compared to samples of the same genus from other localities of the San Pedro Valley of comparable age, and some inferences on the taxonomy of the genus are attempted. The specimens are referred to Sigmodon minor and Sigmodon cf. S. curtisi. Other cricetids (Onychomys pedroensis and Baiomys brachygnathus) and arvicolids (Mictomys vetus and Ondatra ídahoensis) are also recognized and described. 

The study of that locality allowed the description of a new Muridae : Paraethomys athmeniae n. sp. It reveals the existence of new rodent for Algeria : first, a Sciuridae, Atlantoxerus cf. rhodius, and second, a Gliridae, Eliomys truci. So, that work shows the presence of the genus Eliomys in North Africa before the middle of Pleistocene. Lastly, Paraethomys cf. anomalus gives an exact datation of that bed. 

The comparative morphology and biometry of the lower cheek teeth of modern Equus are studied on approximately 300 mandibles belonging to the 10 usually recognised species : Equus grevyi, E. burchelli, E. quagga, E. zebra, E. africanus, E. asinus, E. hemionus, E. klang, E. przewalskii, E. caballus. The studied parameters comprise : occlusal length and width, postflexid length and index ; shape of the double knot (metaconid + metastylid + lingual groove) ; depth of the vestibular groove on the molars ; frequency of the pli caballinid, protostylid and other enamel plications or islets ; frequency of the dP/l.
The same methods of study are applied to a number of North American, Eurasian and African species. For the sake of comparison, some Hemphillian equids were observed (Dinohippus interpolatus, Dinohippus leidyanus, Astrohippus ansae, Phiohippus mexicanus) but most of the discussed material belongs to Pliocene or Pleistocene species of Equus : the « stenonine ›› E. stenonis, E. simplicidens, E. sanmeniensis and E. teilhardi; the « caballine ›› E. scotti, E. lambei, E. Iaurentius, E. mosbachensis, E. germanicus, E. gallicus, E. taubachensis and the Liakhov horse. The relationships of other species, in particular the North American E. calobatus, E. occidentalis, E. cf mexicanus are not clear for the moment. ln Africa, the Plio-Pleistocene species from Koobi Fora (Kenya) show some stenonine and perhaps asinine affinities. The relationships of E. numidicus and E. tabeti are uncertain but these species are probably related to the East African ones. E. mauritanicus is most certainly related to the Quagga group.
The biometrical data are gathered in 32 tables ; 4 photographie plates and 19 figures illustrate the next. The whole is a complement of the previously published studies of the skulls, upper cheek teeth, incisors and metapodials of modern and fossil Equus.

  

The Pantolestidae from the middle eocene of Bouxwiller are the subject of a detailed study. Buxolestes hammeli (n. g., n. sp.) is not closely related to any other European or North American form described until now; it presents, however, some characters in common with Pantolestes, a form of the same age from North America. A parallel evolution from a common ancestral form could explain this ressemblance.
Another form (gen. and sp. indet.) accompanies Buxolertes hammeli in the Bouxwiller fauna.
  

The genus Microstonyx was found only in the north eastern part of Spain : M. antiquus, referable to the
Eppelsheim type, from the pyrenean basins ; M. antiquus, more progresive, from the Vallès Basin : M. major-erymanthius group, from the Teruel-Ademuz Area and from Pieru (Penedès Basin).

  

The mammalian fauna of the Tagai locality of Olhon Island in Lake Baikal contains six artiodactyls:
Amphitragulus boulangeri, Lagomeryx parvulus, Stephanocemas sp., Palaeomeryx kaupi, Orygotherium aff. escheri, and Brachyodus intermedius. They are reported from Eastern Siberia for the first time. A description of teeth and limb-bones are given. An analysis of the systematic composition of the Tagai fauna allows it to be correlated with European faunas of the second part of the Early Miocene (Orleanian, MN 3-4) and permit suggestion of the existance of a united European-Siberian zoogeographical sub-area in the middle latitudes of Eurasia at that time. 

The first lists of Insectivores (Erinaceidae, Talpidae and Soricidae) from the Pliocene beds of Southern France and North-East Spain are given in this paper. The material from twelve localities is studied. These localities are geographically situated in Languedoc (Celleneuve, Vendargues, Nîmes, Sète, Balaruc 2 and Seynes), in Roussillon (Terrats, Serrats-d'en-Vacquer, Château d'eau and Mont-Hélène) and in North-East Spain (Layna, Medas Islands and Puebla de Valverde). These faunas correspond to the Early, Middle and Late Pliocene. 1 to 8 taxa are identified in these localities and 14 specific taxa are presently listed for this period in this area. Two new specific taxa are described as Galerix depereti nov. sp. from all the Early Pliocene localities in the North-Pyrenean area and as Desmanella gardiolensis nov. sp. from Balaruc 2. For this small mammals, two faunal assemblages are recognized. The first one is dated from the Early Pliocene (F 1, 2 and 3 zones in Aguilar et Michaux) and is characterized by Galerix depereti and rare and little diversified Soricids. The second one is Late Pliocene in age (zones G 2 and G 3). The fossils of the genus Talpa are relatively abundant and the Soricids are diversified and very abundant. The Middle Pliocene (zone G 1) is a transitional period. ln these faunas, most of the insectivore genera are known from the European Late Miocene beds (8 on 10). This fact demonstrates a relative continuity between the invectivore faunas from the Late Miocene to the Early Pliocene. In conclusion, somme paleoecological considerations are suggested.
  

This study describes four taxa of Gliridae from the Oligocene mammal locality Gröben 3: Gliravus tenuis BAI-ILO, 1975, Bransatoglis micio (MISONNE, 1957), B. planus (BAHLO, 1975) and B. heissigi n. sp. Gliravus tenuis from Gröben 3 is somewhat more advanced than the type population found in Heimersheim. This confirms previous research suggesting that Gröben 3 should be dated earlier than Heimersheim (MP 24). The first documented occurrence of B. mício around level MP 24 was found in Gröben 3. An abundance of tooth material from B. planus in Gröben 3 makes it possible, for the first time, to observe evolutionary stages within this species from MP 21 until MP 28. B. heissigi n. sp. is restricted to level MP 24. This species is located between B. mísonnei (MP 20 - 23) and Microdyromys praemurinus (MP 25 - 28). Within the lineage Bransatoglis bahloi - B. misonnei - B. heissigi, a decrease in size is noticeable.

  

The principles and practices employed in establishment and recognition of South American land mammal ages are reviewed along with previous and present concepts of distinguishing time, rock, and faunal units. Previous chronological arrangements of South American Tertiary land mammal faunas are appraised on the basis of recent geological and paleontological data. Twelve South American Tertiary land mammal ages are here recognized [from oldest to youngest, Riochican (middle to late Paleocene); Casamayoran (early Eocene); Mustersan (middle Eocene); Divisaderan (late Eocene); Deseadan (early [to middle?] Oligocene); Colhuehuapian (late Oligocene); Santacrucian (early Miocene); Friasan (middle Miocene); Chasicoan (late Miocene); Huayquerian (latest Miocene); Montehermosan (early to middle Pliocene); and Chapadmalalan (late Pliocene)]. As all except the Friasian were originally defined on the basis of Argentine faunas, these are discussed first and at length, and each is reviewed with discussion of type locality, stratigraphy, type fauna, and faunal correlations. Non-Argentine faunas are then discussed country by country in alphabetical order.

     A review is given of radioisotope dates obtained on volcanic rocks (i.e., basalts, tuffs) associated with mammalbearing beds in Argentina. Based on these age determinations and on correlation of the late Tertiary land mammals involved in the interchange between North and South America, a chronology of South American land mammal ages correlated with North American land mammal ages and European marine stages is proposed.

It is concluded that South America was an island continent through most of the Tertiary Period (ca 65 to about 3 Ma). As a result, the land mammal fauna of South America developed in isolation and was dominated by autochthonous endemic groups. Toward the end of the Tertiary (i.e., middle Miocene) a unique faunal balance had been achieved by the descendants of the ancient inhabitants (notoungulates, litopterns, condylarths, astrapotheres, edentates, marsupials) and of later (late Eocene) waif immigrants (caviomorph rodents, platyrrhine primates). A prominent feature of this mammal fauna was the combination of carnivorous and omnivorous marsupials with native placental herbivorous ungulates, subungulates, and edentates.

Sometime during the late Miocene, a limited but important interchange of mammalian taxa between North and South America took place. Procyonids (raccoons and their allies), a group of North American origin, first appear in South America in strata of Huayquerian Age, while members of the extinct South American ground sloth families Megalonychidae and Mylodontidae first appear in North America in early Hemphillian time. These groups dispersed along island arcs before the appearance of the Panamanian land bridge in the Pliocene (ca 3.0 Ma). Cricetine rodents, a group of North American origin, are first known in South America in strata of Montehermosan Age. The known taxa are too advanced and diversified to be considered the first of this group to invade South America. lt is believed by some workers that these rodents arrived before the Montehermosan, possibly in the late Miocene or earlier, by waif dispersal from North America.

The isolation of South America ended with the appearance of the Panamanian land bridge, which provided a direct, dry land connection between the two Americas. Across this portal an extensive interchange of terrestrial faunas occurred, and the fossil record documents an intermingling of these long-separated land mammals faunas.

      The beginning of this interchange by land route in South America is marked by the appearance of mammals which evolved from North American emigrants in the Chapadmalal Formation of Argentina. These include a mustelid (Conepatus), a tayassuid (Argyrohyus), and four genera (Akodon, Dankomys, Graomys, Reithrodon) of cricetine rodents. The appearance of this contingent of northern animals favors the existence of the Panamanian land bridge by this time. Likewise, a large number of terrestrial vertebrates of South American origin appear in North America in beds of late Blancan Age date around 2.7 Ma. Among the mammals are Neochoerus, Erethizon, Glyptotherium, Glossotherium, Kraglievichia, and Dasypus.