The Late Miocene site of Gritsev (MN 9, Ukraine) has yielded a very rich bat fauna, the remains of which are well preserved. Compared to other Neogene bat assemblages of Europe, the Gritsev bat community is unique in preserving plecotine bats, which are rare from Neogene sites. Some peculiar and new bat species, including a large plecotin Otonycteris, already were described from the Gritsev mammal site. Here we report new records of small plecotin bats from Gritsev, including a new taxon, Barbastella maxima nov. sp. This is the earliest reliable fossil record of this genus and it differs from more recent species of Barbastella in being considerably larger. The evolutionary patterns in the odontology within the tribe Plecotini, supported by biostratigraphical distribution of fossil records of Plecotus are discussed. The morphological peculiarities of the new fossils of plecotine bats from Gritsev are discussed in connection with its possible taxonomical affinity. 

The early gnathostome genus Mesacanthus is well represented in both Lower Old Red Sandstone and Middle Old Red Sandstone assemblages of northern and central Scotland. This ‘acanthodian’ taxon is currently thought to comprise two valid species: M. mitchelli and M. pusillus. Although the whereabouts of the holotype of M. mitchelli (NHMUK PV P560) is known, the syntype material for M. pusillus has long been thought lost. Here we identify at least one specimen that formed part of the original syntype material for M. pusillus, albeit in a slightly different condition than when it was originally figured. This specimen is ROM 25872, which is here designated as the lectotype. A second specimen – ELGNM 1978.191.1 – could represent another of the syntype specimens, but poor preservation quality makes it impossible to be certain. 

Abstract: Paleontological exploration in the Turkana Basin near Ileret, Kenya yielded the most complete adult elephant cranium (KNM-ER 63642) known from the late Miocene to mid-Pliocene. KNM-ER 63642 derives from the lower Lonyumun Mb. of the Koobi Fora Fm. and dates to the early Pliocene, >4.3 Ma. The cranium is immense in size and preserves most of its structures including left and right M2-3, permitting its comprehensive comparative study and secure taxonomic assignment to Loxodonta adaurora. Features distinctive of the species and exhibited by KNM-ER 63642 include very elongate, divergent tusk alveoli, a short, biconvex cranial roof, anterosuperior angulation of the occipital planum, non-inflated occipital planum and absence of supralateral parietal "bossing," broad, flat premaxillary nasal processes, broad, laterally downturned nasal aperture superior to the level of the orbits, and M3s with wide, subhypsodont plates that are parallel-faced and separated by U-shaped transverse valleys. The M3s also exhibit characteristic L. adaurora traits of greatest width at their bases, rounded cross-sectional shape, thick enamel, abundant cementum, and strong anterior and posterior accessory conules. Of extant taxa, KNM-ER 63642 most closely resembles crania of African elephants. Its inclusion in the Loxodonta clade is tenuous, however, because shared features are either symplesiomorphic or are difficult to test for synapomorphy due to the poor fossil record of crania of late Miocene-early Pliocene elephants. Overall, the cranial morphology of KNM-ER 63642 is unexpectedly advanced for an elephant of its antiquity. Its anteroposterior compression and height are concordant with efficient proal masticatory action, indicating that by the early Pliocene L. adaurora evolved craniodental adaptations in phase with feeding preference for C4 grasses. The advantage of synchrony of morphology and behavior is reflected by the dominance of the species in the greater Turkana Basin during that interval.
  

Record of life and works of Henri Menu, French zoologist, contributor to the knowledge of living and fossil bats. 

Evidence of 350 Lower Cretaceous Dinosaur footprints is pointed out in El Bayadh area. Their preliminary study allow to distinguish four trackway assemblages which reveal vertebrate bipedal presence forms of tri-and tetradactylous Dinosauroïds (Assemblages 1-3) and quadrupidal Sauropoïd (Assemblage 4).

The analysis of their footprint biometric features will attribute the quadrupidal Sauropoïd form to Brontopodus ichnogenus which is weIl known in the Jurassic and Cretaceous periods. In retum and despite their age, the dinosauroïd forms were approached, temporarily, to Grallator and Eubrontes types.

The occurrence of the dinosaur traces (Theropoïd and Sauropoïd) constitutes, in the Lower Cretaceous, an important first step of the knowlege of the marshy Reptilian fauna which takes over, from the begining of the Secondary Era, a wide paleogeographie area on the Southem Tethyan margin. 

Since the 19th century, thousands of footprints were observed in the geological series of the French Mesozoic. All are located in the Triassic and Jurassic. After a promising beginning, in France, it is only a few papers which will be published in the first half 20th century, unlike the USA and of others countries of Western Europe. One ought to wait about 1950 for a revival and now they are nearly 200 papers which were devoted to the ichnofossils. The literature abundance and the renewed interest of the naturalists for the palichnologic studies decided to us to write a synthesis work. This one begins with a stratigraphic inventory in which, localisation, age and paleontological contents of about 180 fossiliferous sites are specified. After having pointed out the followed methods, the footprints paleontological interpretation is then approached in detail and the results obtained are replaced in stratigraphy to deduce the fauna evolution during the Mesozoic. So, it appears that Ichnologic data, more varied and rich in the Triassic and Liassic than those relating to the bones, very rare for the considered periods, are very informative. The middle Triassic (Anisian-Ladinian), thus reveals Cotylosauria, Lepidosauria, Crurotarsi with Rauisuchia, Ornithosuchidae, Crocodylia and Dinosauromorpha more the "Prodinosauria": Dinosamiforme whose skeletons are known in Argentina but only in Ladinian. The rather fast domination of Dinosaurs during Norian is also as well shown. The almost exclusive presence of their footprints, up to fifty cm long, in the Lower Hettangian indicates their supremacy in the environments. Footprints characterise not very deep life places located between inter-supratidal limits and often out of water. Sedimentologic and Palaeontologic studies showed that they were great coastal spaces during Middle Triassic, flood-plain with sebkhas while Upper Triassic, and a large !!coastal marsh!! in Grands-Causses during Liassic in which, mainly, fine stromatolithic layers were deposited. During the same periad, bay beaches spread in Vendée. During the Middle Jurassic, they are also brackish to lacustrine environments and recifallagoons in- the Upper Jurassic. Numerous measurements of the footprints and trackways directions showed that the animaIs moved there in weil defined directions, for long periods. They seem due to the palaeotopography of the life environments relatively stable. Also, the discovery of vegetal radicular networks and small footprints far away from the continental borderlands has suggested that the animals continuously lived in these palaeoenvironnements, belonging to large ecosystems, where the sedimentation rate was weak. This explains that thebadies could not fossilize there but only their footprints through the cyanobacterian action in main cases. From the vertical distribution of different ichnospecies, defined with adapted statistical methods, explained in this work, a palichnostratigraphy was established for the Middle Triassic. Although the footprints are also abundant in Hettango-Sinemurian of "Grands-Causses" and the Vendée, it was not possible, up to now, to establish any zonation in this series; Probably because the palichnofauna is too little diversified there, currently reduced to a majority of Theropods II-IV tridactyl traces.
  

A new carcharhinid shark species, Physogaleus hemmooriensis sp. nov., is described from the Lower Hemmoorian (Behrendorfian, late Burdigalian, early Miocene) of Werder, Lower Saxony, Germany. P. hemmooriensis also occurs in the Edegem and Antwerpen Sands Members of the Berchem Formation, Belgium, and in the Miste Bed, Aalten Member of the Breda Formation, The Netherlands, which have an early to middle Miocene age. In the Western Atlantic region, the taxon is present in the early Miocene Calvert Formation of Delaware, U.S.A, which is largely contemporaneous with the Hemmoorian. 

In recent years, the techniques of chemical preparing have permitted a rich paleontologic material to be obtained from the phosporitic sediment of Bouzigues (Hérault, France). The fauna of this locality is comprised of quite varied microvertebrates, amphibians, reptiles, birds, mammals. Twenty five species of the latter, belonging to seven orders, are today known from the site. Among them, the numerous rodents have allowed L. Thaler to chronologically situate this fauna in the Zone of Laugnac (<< late Aquitanian ›> of some authors).
The chiropterans are, with the rodents, the best represented of the locality's mammals. Three families comprise the bat fauna, with nearly complete dominance by one of them (Hippoxideridae) over the two others (Megadermatidae and Vespertílionidae)
Six forms are described, of which three are new species and one a new sub-genus.
Megaderma braillomi n. sp., an animal of rather large size, shows like the Miocene megaderms several evolved dental characters, translating the adaptation of these animals to a partially carnivorous regime. The Bouzigues species seems, however, to represent a particular lineage.
Hipposideros (Brachipposideros n. subgen.) dechaseauxi n. sp. and Hípposideros (Brachipposideros) cf. collongenris Depéret, small sized forms, belong to a group rather well represented in the late Oligocene and early Miocene of Europe, and not distinguished until now within the genus Hipposideros.
Hipposideros (Pseudorhinolophus) bouziguensis n. sp., is the most abundant mammal in the locality and, occuring at the Oligocene-Miocene limit, the last representative known of the subgenus Pseudorhínolophus, common in Europe from the middle Eocene.
However, beyond Neogene and Quaternary times, certain among the numerous living species of Hipposideros are close to Pseudorhinolophus and others to Brachipposíderos. 'This fact would in the future justify a global revision of the genus, on the basis of comparative anatomy of the squeleton and of the teeth.
The bat fauna of Bouzigues is completed by two small Vespertilionidae, rare forms, Myoris sp. I and sp. II.
  

A late Pleistocene deposit of 60 species of vertebrates and 12 of invertebrates is described from Robinson Cave, Overton County, Tennessee, U.S.A. Forty-eight species of mammals are represented by at least 2,483 individuals; 10 % are extinct, 10 % occur in the state only as boreal relicts in the Great Smoky Mountains; 23 % no longer occur as far south as Tennessee; 57 % occur at or near the site today. Nínety-one percent of the Recent mammal species can be found living today in the Minnesota-Wisconsin area, approximately 10 degrees farther north. Fluorine analysis suggests a long period of accumulation. The following 10 mammalian species are recorded from Tennessee for the first time. Sorex arcticus, Microsorex hoyi, Citellus tridecemlineatus, Clethrionomys gapperi, Microtus pennsylvanicus, Synaptomys cooperi, Synaptomys borealis, Zapus nudsonius, Napaeozapus insignis, Martes americana. Six additional species are present as boreal relicts in the Great Smoky Mountains of eastern Tennessee but not at the site today : Sorex cinereus, Sorex dispar, Sorex palustris, Parascalops breweri, Glaucomys sabrinus, Mustela nivalis. Six forms are extinct: Canis dirus, Ursus americanus amplidens, Sangamona furtiva, Dasypus bellus, Mammut americanus,Megalonyx jeffersoni. Twenty-six additional species of mammals, all of the snails, birds, reptiles, and amphibians recovered from the fauna still inhabit the area today: The fauna is indicative of a cold-temperate climatic episode associated with the Wisconsin glaciation, but may be chronologically mixed. 

Description of four new rodents from a recently discovered locality at Montalban. Theridomys crusafonti nov. sp. is considered as the ancestry of T. Iembronicus. Theridomys varian: nov. sp. includes «Theridomys» morphotypes and «Blainvilllimys» morphotypes; it could be ancestral to B. blainvillei. Pseudoltinomys nanus nov. sp. represents a new lineage paralleling in evolution that of P. gaillardi (which is equally found at Montalban). Pseudocricetodon montalbanensis nov. gen., nov. sp. designates a lineage of very small Cricetidae accompanying Eucricetodon. With these well defined new species and six others present in the locality, Montalban appears as the best faunal reference point within the biochronologic zone of La Sauvetat.
As an annex, discussion of two rodent specimens from the classic localíty of Tárrega, close in age to that of Montalban. 

Based on isolated teeth two new scyliorhinid shark species, Scyliorhinus biformis nov. sp. and Scyliorhinus suelstorfensis nov. sp., are described from the Sülstorf Beds, early-middle Chattian, of Mecklenburg, northeastern Germany. They form part of a speciose assemblage of necto-benthic sharks and batoids which populated the warm-temperate to subtropical upper shelf sea of the south-eastern North Sea Basin. 

Based on fossil discoveries and phylogenetic studies, an Eocene Asian origin for hystricognathous rodents and anthropoid primates has gained strong support in recent years. The two groups then invaded both Africa and South America, which promoted their evolutionary success. However, the fossil record has so far failed to constrain the nature and precise timing of these pivotal dispersal events. In Africa, given the apparent absence of hystricognaths and anthropoids in early to early middle Eocene localities, it is suggested that these mammal groups dispersed from Asia to Africa sometime during the middle Eocene. In this paper, we report the discovery of several isolated teeth of a rodent from a new vertebrate locality situated in central Tunisia (Djebel el Kébar, KEB-1), dating from the late middle Eocene (Bartonian, ~39.5 Myr). These fossils document a diminutive new species of Protophiomys (P. tunisiensis nov. sp.), a basal genus of hystricognathous rodents which is well known from several North African mammalian-bearing localities of the end of the Eocene. The teeth of P. tunisiensis display a suite of anatomical details comparable with those observed in the other species of the genus, but with a lesser degree of development. Such an apparent primitive evolutionary stage is corroborated by the greater antiquity of this Tunisian species. P. tunisiensis nov. sp. is so far the most ancient representative of hystricognaths in Africa. However, it can be expected that hystricognaths were already present on that landmass given the new data on early caviomorphs recently reported from South America (at ~41 Myr). The arrival of hystricognaths in Africa from South Asia certainly predates the depositional period of the Kébar sediments, but perhaps not by much time. 

The Oligocene deposits of the Ulantatal area in Inner Mongolia (China) contain among the richest mammalian faunas from Asia. To date, only some parts of the rodent faunas have been described. Here, we propose to review the rodent faunal lists for each site, including the description of a few new rodent specimens. We describe three additional rodent species: the Cylindrodontidae Anomoemys lohiculus, the Eomyidae Asianeomys sp., and the Dipodidae Litodonomys huangheensis. This study allows us to constrain the stratigraphic range of Anomoemys lohiculus, which ranged from the late Early Oligocene to the early Late Oligocene in this area. Asianeomys sp. and Litodonomys huangheensis are dated from the latest Oligocene. These Oligocene deposits consist now of more than 70 species of mammals if we include the fauna from Kekeamu. This latter corresponds to the basal part of the Ulantatal Formation and could be dated biochronologically from the earliest Oligocene. When compared to the faunas from the Valley of Lakes in Central Mongolia, the Ulantatal faunas present a great majority of rodents, and this difference can be partly explained by sampling and description biases regarding macro-mammals. This study also shows that variations existed between Inner and Central Mongolia, especially regarding the composition of the rodent paleocommunities. However, the assessment of their evolutionary history in this part of Asia with respect to the important climate and environment changes, require further precisions and more material than current data allow. 

A comparative bone histological study of the three known genera of the endemic European ornithopod dinosaur family, Rhabdodontidae, is presented here in an ontogenetic context. Investigated specimens were assigned to different ontogenetic stages based exclusively on the histological indicators of osteologic maturation during diametrical bone growth; an entirely size-independent method as opposed to most previous studies. Qualitative comparison of bone histology of corresponding ontogenetic stages and elements among the three valid rhabdodontid genera, Mochlodon, Zalmoxes, and Rhabdodon, revealed some consistent patterns. Genus specific histological differences within Rhabdodontidae are most expressed between Rhabdodon and the Mochlodon-Zalmoxes clade. These indicate a prolonged phase of fast growth and a less constrained cyclicity in the growth dynamics of Rhabdodon, as opposed to the slower and more regulated growth strategy reflected in the bones of Mochlodon and Zalmoxes. These genus specific differences are consistent with the phylogenetic interrelation of the genera and are most probably related to the pronounced differences in body size. However, when compared to other ornithopods, most detected histological features in rhabdodontids do not seem to reliably reflect either phylogenetic relations or body size. A notable common feature of all rhabdodontid genera irrespective of body size is the ontogenetically early onset of cyclical growth and secondary remodelling; a pattern that more resembles the condition found in derived ornithopods than that described in more basal taxa which are closer relatives of rhabdodontids. The recognition of taxon-specific histological patterns as well as patterns indicative of ecological and thereby functional traits clearly requires more accurate, preferably quantitative evaluations.   

The earliest cetaceans are found in the early Eocene of Indo-Pakistan. By the late middle to late Eocene, the group colonized most oceans of the planet. This late Eocene worldwide distribution clearly indicates that their dispersal took place during the middle Eocene (Lutetian). We report here the first discovery of a protocetid fossil from middle Eocene deposits of Senegal (West Africa). The Lutetian cetacean specimen from Senegal is a partial left innominate. Its overall form and proportions, particularly the well-formed lunate surface with a deep and narrow acetabular notch, and the complete absence of pachyostosis and osteosclerosis, mark it as a probable middle Eocene protocetid cetacean. Its size corresponds to the newly described Togocetus traversei from the Lutetian deposits of Togo. However, no innominate is known for the Togolese protocetid, which precludes any direct comparison between the two West African sites. The Senegalese innominate documents a new early occurrence of this marine group in West Africa and supports an early dispersal of these aquatic mammals by the middle Eocene.
  

Nyctereutes megamastoides (Pomel) from the Villafranchian of the Auvergne and from Saint-Vallier presents cranial and dental characters sufficiently close to those of the late Pliocene canid from Perpignan (Roussillon), described by Depéret under the specific name of Canis donnezani belonging to the same genus Nyctereutes. The extinction of the European "Nyctereutes" group seems due to the too great alimentary specialization of this canid, whereas the Asiatic lineage represented in the Villafranchian by Nyctereutes sinensis Schlosser and at present by Nyctereutes procyonider Gray was able to maintain itself probably by means of a profound change in its alimentary regime. 

The scientific career of D.E. Russell began with a Pliocene fauna from Oregon, and then turned in the direction of European Paleogene mammals. Field work followed by study of the mammals that were collected, firstly in the Paleocene and later in the early Eocene, greatly rejuvenated learning in this field. Syntheses on the Northwest European Tertiary basin and on European marnmals and stratigraphy came next. Research on the Eocene of Asia was carried out jointly with Gingerich on Pakistan and with Dashzeveg on the faunas of Mongolia. An important synthesis on the entire Paleogene of Asia, joint with Zhai, followed. Field work in Africa with Sigogneau-Russell led to the discovery of Mesozoic mammals there. A synthesis of mammalian paleofaunas of the world was written with Savage, and a similar synthesis of Cenozoic vertebrate faunas is currently being prepared. These achievements reflect the perennial importance of field work, numerous collaborations with both amateurs and professionals, and the human qualities of this author.
  

Le présent volume est l'aboutissement d'un projet né il y a presque cinq ans. En décembre 1991, l'un d'entre nous (MG) prenait des contacts en vue de proposer un symposium sur les mammifères fossiles, dédié à D.E. Russell, dans le programme du 4e Congrès de la European Society for Evolutionary Biology. Ce congrès, baptisé "Evolution 93", devait se tenir à Montpellier en août 1993. Son Comité d'Organisation, animé par F. Catzeflis, recherchait des organisateurs de symposiums. L'idée fut acceptée avec enthousiasme par le second d'entre nous (PDG), et le titre de notre Symposium fut précisé: " Palaeobiology and Evolution of Early Cenozoic Mammals - A Symposium in Honor of D.E. Russell". Le projet fut formellement accepté par le Comité d'Organisation en avril 1992. 

The rodent-fauna (Cricetidae and Gliridae) recently found at Castell de Barbera (Spain) is similar to those from the other locslities of the Valles - Penedes - Can Ponsic 1 and Can Llobateres - : same composition and similar evolutionary level of the different species. On the other hand this fauna is different from those of Upper Vindobonian and Vallesian localities of the Calatayud - Teruel area. Castell de Barbera has an intermediate chronological position between the localities of Anwil (Switzerland) and Can Ponsic 1. It is still not possible to validate or invalidate the initial attribution based on absence of Hipparion of Castell de Barbera to the Upper Vindobonian. 

Until now Aplodontidae of the European Oligocene have been documented by four species only. The phylogenetic relations remained obscure. as the distribution of only one species has been known in some detail. New material made it possible to define the stratigraphic range of two of the already existing species (Plesispermophilus angustidens, Sciurodon cadurcense) and to follow their development during the Oligocene beginning with the event of the « Grande Coupure ››. Sciurodon remained nearly without change until the end of the Middle Oligocene. Plesispermophilus angustidens split into two distinct phyletic lines, one of which (P. macrodon n. sp.) reaching considerable size, is represented till the beginning of the Upper Oligocene (Pech de Fraysse, Gaimersheim). The other line leads to Plesispermophilus ernii (basal Upper Oligocene of Burgmagerbein 1. terminal Upper Oligocene of Coderet). Besides the already known forms a new small-sized species (P. atavus n. sp.) is described, which by its primitive features closely resembles the genus Plesispermophilus. Two other small-sized species already known from the Upper Oligocene (? P. argoviensis) and Lower Miocene (? P. descedens) seem to be closely related to the new species. It cannot be decided whether they are descendents of this line or have developed independently, because of their poor fossil record.
Comparison of the evolutionary modalities in the different phylogenetic lines reveals general trends. the most striking of which is the complication of the tooth pattern by the development of additional crests. In the lower molars the cusps diminuate in size and are more and more transformed into ridges. ln addition new connection between the crests appear. in the upper molars, the « selenodont » shape of the teeth becomes more and more dominant, and in the two main evolutionary lines of Plesispermophilus the metaconulus becomes duplicated. A further evolutionary trend is the size increase of the premolars compared to the molars, which is even more pronounced in the Miocene Aplodontidae.
Phylogenetic relations between the primitive Plesispermophilus and certain « prociurines ›› of Northern America as well as between Plesispermophilus (P. angustidens) and more progressive forms of the Upper Oligocene (P. ernii, P. macrodon n. sp.) can be documented. In this light, the taxonomic distinction between Prosciurinae (bunodont) and Allomyinae (selenodont) sensu Rensberger 1976 can be shown to be artificial, because it separates forms from each other, which are evidently closely related. Consequently the separation into two subfamilies has been abolished.