Anthracotheres from the Pondaung Formation, Myanmar, are considered as one of the most primitive artiodactyl groups and they represent the oldest known record in the world. Thus, the understanding of this group has numerous implications for evolutionary biology and biochronological correlations. However, the systematlcs of these mammals has been interpreted in different ways, and the main debate focuses on the number of taxa represented in the Pondaung Formation. The revised taxonomy proposed here is mainly based on the relative development of the upper molar W-shaped ectoloph, system of crests and stylar cusps, and on body size. On the basis of these characters, they are classified into four genera including six different species. Two well-known genera, Anthracotherium and Anthracokeryx, are validated and more precisely diagnosed. Anthracokeryx possesses a better developed W-shaped ectoloph, system of crests and stylar cusps than Anthracotherium, which displays notable differences with the more derived representatives of this genus. Both of these Pondaung genera show evidence for sexual dimorphism. However, the incompleteness of fossil material fueled a debate concerning the status of two additional Pondaung anthracotheres, Siamotherium and Anthracohyus. The latter genus is of uncertain affinities, but it has been considered as a hippopotamid ancestor. Despite new material attributed to these two forms, additional discoveries are still required to establish their taxonomic status. The hypothesis that Southeast Asia was the centre of origin of Anthracotheriidae is supported by the retention of numerous primitive dental characters in these taxa and by the antiquity of the Pondaung Formation, to which an age of 37 My is now generally accepted. 

A fragmentary mandible and maxilla of a small sized Agriotherium of a young individual discovered from a Pliocene fissure filling in Xiaoxian county (Anhuei Province, China) are described. Judging from the morphology of the dentition and its dimensions the new material can be identified as Agriotherium inlermedium (STACH l957). Hendey's proposition (1980) that the Agriotherium species are derived from Indarctos is reconsídered on the basis of the new documents. As a result of a more general phylogenetic discussion it can be stated, that: 1. the supposed size increase as well as other trends, leading from Indarctos to Agriotherium are untenable ; 2. there are no positive indications to assume a phylogenetic transition of these two genera. 3. there are no real arguments in favor of an adaptational reversal in the evolution of Agriotherium. Hence, many features of that genus supposed by Hendey to be derived are plesiomorphic ; 4. regardless of the previous points it is methodologícally impossible to establish direct ancestor - descendant relationships between Indarctos and Agriotherium species, as Hendey did. Based on the data available and especially on the characters of the new material from China it is more likely that Agriotherium and Indarctos are two genera which developed independently. While advanced Agriotherium species, e.g. A. africanum, resemble in some respects Indarctos by adaptational analogies, more primitive species, e.g. Agriotherium intermedium, are quite dissimilar to lndarctos. While Indarctos might be derived from an Ursavus like forerunner, Agriotherium has its roots more likely somewhere in between Ursavus and the Hemicyon-group. 

Enamel microstructure in the incisors of Old- and New World hystricognath rodents:

The incisor enamel microstructure in more than 100 genera of fossil and Recent hystricognath and sciurognath rodents was studied. A multiserial schmelzmuster is present in the Hystricognathi, the Ctenodactylidae, advanced Chapattimyidae, and in Pedetes. A redefinition of pauciserial and multiserial HSB is given that makes the two enamel types unambiguously distinguishable which apparently represent well defined evolutionary levels. In the pauciserial Schmelzmuster the IPM is thicker than in the multiserial one. In pauciserial HSB the IPM always surrounds each prism, and the crystallites of the IPM run parallel to prism direction; transition zones between HSB are lacking; the inclination of the HSB is normally very low and the prism cross sections are not flattened but somewhat irregular. The number of prisms per HSB is no good distinctive character for pauciserial and multiserial HSB, since there exists a wide overlap. The pauciserial schmelzmuster is primitive, the multiseiial derived because: 1. the pauciseiial schmelzmuster appears earlier in the fossil record in the most primitive rodents (Paramyids s.l. and Ctenodactyloids); 2. the Eocene Ctenodactyloidea show pauciserial HSB but the Oligocene and younger ones are characterized by multiserial HSB; 3. in the outgroup comparison, the Eurymylidae (Mixodontia) show pauciserial HSB; 4. biomechanically, multiserial HSB strenghten the enamel better than pauciserial HSB, since their IPM runs nearly always in an angle of 45° or more to the prisms.

In multiseríal HSB three subtypes can be distinguished which are differentiated by the IPM orientation. Primitive is a (rarely strict) parallel or acute angular, anastomozing IPM, and derived is an interrow sheet-like ("plattenartige") IPM. This evolutionary polarity is indicated by enamel evolution in the Ctenodactylidae which show an acute angular IPM in the Oligocene and a rectangular interrow sheet-like IPM since the Miocene. Among the Caviomorpha a rectangular interrow sheet-like IPM is restricted to the Octodontoidea; therefore they must be considered derived in terms of their enamel structure. The first multiserial HSB in rodent incisors appear in phiomyids or chapatrimyids from the Upper Eocene of Algeria. The IPM is acute angular and anastomozing. The worldwide next younger multiserial HSB are found in Lower Oligocene phiomyids of Fayum, Egypt There already a rectangular interrow sheet like IPM is present (in Metaphiomys) besides the acute angular anastomozing IPM.

The first Caviomorpha from the Deseadan (Oligocene-Miocene) likewise show already acute angular anastomozing IPM (e.g. Scozamys) and rectangular interrow sheet-like IPM (Platypittamys). Therefore the first Caviomorpha cannot be positioned close to a transition from pauciserial to multiserial HSB. In none of the potential caviomorph ancestors from southern North America multiserial HSB or transitional stage between pauciserial and multiserial HSB could be found. The similarities between the enamel types of the Fayum rodents and the rodents from the Deseadan of South America make a derivation of the Caviomorpha from Paleogene North African phiomorph rodents or their direct ancestors most probable. This supports at the same time a descent of the platyrrhine Primates from North African anthropoids.
  

Cet article présente une étude détaillée du genre Stehlinia, un chiroptère du Paléogène d'Europe, basé sur un matériel abondant issu notamment du gisement d'Escamps (Quercy). L'analyse révèle que Stehlinia possède un mélange de caractères primitifs (comme une denture tribosphénique complète et des prémaxillaires soudés) et évolués (notamment dans le squelette post-crânien), le rapprochant des vespertilionoïdes actuels, en particulier des Kerivoula. 

An isolated right costal 1 from the Late Cretaceous Massecaps locality (Cruzy, Hérault, southern France) is assigned to Dortoka vasconica (Dortokidae). This find adds a new element to the Late Cretaceous turtle fauna of Cruzy and further supports the hypothesis that two distinct lineages of Dortokidae were present in Europe during the Late Cretaceous-Paleogene due to geographical isolation.
  

 A new genus of hystricognath rodent with two new species, Soriamys gaimanensis and Soriamys ganganensis, from the Colhuehuapian Age (Early Miocene), Sarmiento Formation, of Patagonia is described. The first species comes from the south barranca of the valley of Rio Chubut, near Gaiman locality (Chubut Province). It is known through a great number of dental remains in different stages of wear. The teeth are protohypsodont, with cement in the principal valleys. P4 are simpler, more oval inoutline and without a differentiated hypocone-hypoflexus. Upper molars are pentalophodont in early stages of wear and become bilobated in middle and advanced stages; M3 have a temporary third posterior lobe due to the maintenance of the braquiflexus on the posterior wall of the tooth. Lower teeth have a completely asymrnetrical pattern in relation to the uppers. The molars are trilophodont, with only one complex crest in the trigonid composed of the fusion of the anterolophid and mesolophid. The dp4 have a derived pentalophodont pattern due to the loss or fusion of the mesolophid and the development of an accessory transverse crest between the anterolophid and metalophid. The second species is known through a complete skull and jaw and other skull fragments with the dentition, proceeding from Pampa de Gan Gan, Chubut province. Respect to the first species, the teeth are higher crowned, with an earlier occlusal simplification and a third posterior lobe of M3 more developed and persistent. The skull and jaws show many chinchilloid characters, such as jugal bone with an ascending process, concave palate, very prominent dorsal shelf of the masseteric fossa, nasolachrymal duct opened laterally on the maxillary and very developed lateral mandibular fossa. Moreover, Soriamys is closely related to Cephalomys by sharing a similar asymmetric dental pattern and other dental and mandibular characters. These two genera constitute a natural group, the Cephalomyidae, with peculiar dental characters, like the asymmetric pattem of the upper and lower teeth; Cephalomyopsis, and probably Litadontomys, can be considered part of this group. Scotamys and Perimys constitute a distinct lineage (the Neopiblemidae or Perimyidae), more closely related to the chinchillids. Many characters shared by cephalomyids, eocardiids and caviids suggest a closer affinity between the Chinchilloidea and the Cavioiidea as a whole. Abrocoma shows also an asymmetric dental pattern and other chinchilloid features that suggest some degree of relationship with the cephalomyids, still not well determined. 

A fossil fish fauna, based on 5533 otoliths, from the La Paillade locality at Montpellier is described and figured. The otolith-bearing marls correlate to mammal zone MN l (Aguilar, 1982), and thus represent the earliest Miocene. The fish fauna consists of 30 taxa belonging to 20 families. Two species are new: Dussumieria sittigi and Liza gaudanti. The predominant faunal element is the Lesueurigobius vicínalis-species complex, composing 73% of all investigated otoliths. The palaeoecological analysis reveals a marine to euryhaline fish fauna living under tropical to subtropical conditions in the transition zone littoral - sublittoral. Water depth probably was more than 10 m. The scarcity of pelagic físhes suggests that the habitat was either a sheltered bay and/or far away from the open sea. Furthermore, some genera represented in the La Paillade fish fauna presently live exclusively in the Indopacific realm. Their presence strongly supports a broad connection between the Indian Ocean, the Mediterranean, and the Paratethys Seas during the Early Miocene (Aquitanian). From a palaeobiogeographical point of view, faunal relationships were found between the La Paillade fish fauna and both the Paratethys fish fauna and the fish fauna from the deposits in the Upper Rhinegraben and the Mayence and Hanau Basins (Germany).



  

The remains of Ruminantia are very rare at Montredon. No specific determination was possible. We have only one Micromeryx, one small cervid, one tragocere and one (?) gazella. The faunal spectrum is in good correlation with the general retreat of the cervids in the European upper Miocene, but in contrast it is not convenient for the bovids, which are in expansion at the same time. For them, we have to invoke the local ecological conditions.
  

An unpublished mandible of the large hyracoid Titanohyrax andrewsi from the early Oligocene Jebel Qatrani Formation, Fayum Depression, Egypt is described. This specimen has a twofold importance. Firstly, it opens an unexpected window on early paleontological research in the Fayum because it was discovered as early as 1904 by the French paleontologist René Fourtau during an expedition to the Fayum organized by the Muséum National d’Histoire Naturelle, Paris (MNHN). This expedition has remarkably never been mentioned in the literature. Secondly, the mandible documents the best-preserved specimen of T. andrewsi, permitting a revision of one of the very rare Paleogene hyracoids. Interestingly, the new mandible was discovered two years before the first report of the species by Charles W. Andrews. The hypodigm of T. andrewsi is reviewed and the dentition as a whole is compared in detail, notably with other Titanohyrax species from the Fayum. The validity of the large Titanohyrax “schlosseri” species is discussed, but a pronounced sexual size dimorphism for T. andrewsi is favoured. 

The biometric and morphologie variability of the cheek teeth in the end-of-the-phylum species Lophiodon lautricense Noulet studied in this note, reposes on the observation of about 800 teeth. These were revealed to be little variable in absolute dimensions. The considerable morphologie variability in the upper premolars permitted the problem of the molarization process to be taken up. An hypothesis concerning the order of eruption of the cheek teeth is formulated based on an examination of a large number of milk dentitions. In conclusion, it is suggested that reservations be held on the value of dental characters classically used in systematics for the group under consideration. 

A new species, Otonycteris rummeli nov. sp., is described from the Late Miocene site Gritsev (MN 9) in the Ukraine. Otonycteris rummeli nov. sp. differs from those of most vespertilionids, except recent Otonycteris, Antrozous and Early Miocene Karstala silva, in having a well-developed entocingulid at the foot of the trigonid valley in the lower molars. The morphological resemblance of Otonycteris, Antrozous and Karstala is apparently a case of convergence in the evolution of the Old and New Worlds bat faunas. From at least the Middle Miocene the range of Otonycteris distribution spread to the whole of Central Europe and such a situation continued during the whole Late Miocene. This indicates a more arid climate in Europe during the Upper Miocene compared to the Quaternary. The reduction of the distribution range of Otonycteris and its extinction in most of the territory of Europe could have been caused by the global climatic cooling and increasing glacial cycle amplitude during the onset of the Quaternary. 

The rodents (Cricetidae, Gliridae, Sciuridae) found in lacustrine, brackish marine and karstic sediments of Miocene age in Languedoc, assign the position of the different localities in the scale of "niveaux repères" used by mammalogists. Some detailed stratigraphical studies bring several correlations between this continental biochronological scale and the marine scale ; the most important results are the Aquitanian age of the "niveaux repères" of Coderet and Paulhiac, the Burdigalian age of Laugnac, Estrepouy, Vieux-Collonges, La Romieu and Sansan and the Langhian or Lower Serravallian age of La Grive M. The correlations between the Tethys and the Central Paratethys for the Lower Neogene profit also of these results, since the locality of Neudorf Spalte 1, 2 (Czechoslovakia) is shown to be younger than Sansan (France). The paleontological study has also several geological inferences for the Miocene of Languedoc ; with the calibration of this Miocene, we know quite precisely that the Lower Miocene is chiefly a time lacustrine sedimentation, and also that the marine Miocene sedimentation ends early in the Miocene Period, in Langhian or lower Serravallian times. 

Remarkable association of a small infraorbital foramen, similar to that in recent Heterocephalus, and of a strong muscular print on the dorsal anterior part of the zygomatic plate and on the premaxillary. Several anatomical structures to be compared with those of Heterocephalus suggest relationships with this genus. Richardus supports the ancestrality of the hystricomorph character in the bathyergids 

     The Faunas of Rodents from seven north-african fossiliferous beds distributed from the Middle up to the Uppest Miocene are studied. One genus, seventeen species, one subspecies described are new.
       A detailed description is given of the evolution of the numerous specific lines, systematic groups and communities. The evidence of the strong endemism of the North African faunas of that time compared with those of Europe as well as of Tropical Africa is clearly shown. Special relationships with Asia clearly also appear, mostly in the Middle Miocene and the beginning of the Upper Miocene.
       Using the specific lines with a biostratigraphic purpose gives the opportunity to establish a biochronological scale for the continental formations of the Maghreb. This scale is calibrated by a few absolute datations. These datations, as well as the stratigraphical study of several fossiliferous localities have given the possibility to establish precise correlations with the corresponding scales of Europe and of Tropical Africa, as well as with the marine formations of the Neogene of occidental Mediterranean sea.
       The paleoclimatic data connected with this period in North Africa are explained and the contribution of the micromammals is discussed. 

The origin and early evolutionary history of the dinosaurs is a topic that has recently gone through a period of renewed interest and academic debate. For 130 years, one way of classifying the various dinosaur subgroups persisted as the accepted model, with increasing levels of research in the past quarter-century also providing evidence for the hypothesis that dinosaur origination occurred in the Southern Hemisphere, particularly in South America. It is, after all, from within the Late Triassic strata of countries like Argentina and Brazil that we get some of the very best early dinosaur specimens; many of these specimens are the earliest known representatives of some of the major dinosaur subgroups, such as the theropods and sauropodomorphs. However, some recent analyses have brought about a shift in terms of what is currently accepted and what is now disputed regarding the origin of dinosaurs – the Southern Hemisphere origination hypothesis was questioned (although this was based upon observations and not with quantitative analysis techniques), as has the shape of the dinosaur tree. Responses to the new hypothesis were numerous; many further supported a Southern Hemisphere point of origin. Whilst the interrelationships between the major dinosaur clades remains to be resolved, the current data does seem to comprehensively answer the question of where the dinosaurs first originated. However, it is arguable whether the current data that is being used in such palaeobiogeographical analyses is sufficient to provide an answer to the question of where specifically the dinosaur clade first appeared. This short communication urges a degree of caution about the current consensus and what steps may need to be taken to ensure that more meaningful results are produced in the future. 

The dentitions as well as one complete and several partial skeletons of Equoids from the Eocene lignite beds of the Geiseltal locality are revised. Instead of 13 species distinguished up to now 3 chronoclines with 5 species and 3 separate species are recognized (text. fig. 1). Propalaeotherium hassiacum HAUPT, 1925 is evolving into Propalaeotherium isselanum (CUVIER, 1824) between the levels of the « obere Unterkohle ›› and the « untere Mittelkohle ›› of the Geiseltal section. Propalaeotherium argentonicum GERVAIS, 1849 is shown to be present in the « untere Unterkohle ››, whereas Lophiotherium pygmaeum (DEPERET,1901) occurs in the « obere Mittelkohle ›› and in the « oberes Hauptmittel ››. Plagiolophus cartieri STEHLIN, 1904 appears during the transition from the « Mittelkohle ›› into the « Oberkohle ›› as the earliest true Palaeothere. Therefore the « Oberkohle ›› is already regarded as Upper Eocene. This is corroborated by the occurrence of a phyletic descendant of Propalaeatherium parvulum (Propalaeotherium n.sp.) in the middle and upper "Oberkohle " because this species appears otherwise for the first time at the mammal level of Lissieu. On the other hand Propachynolophus gaudryz (LEMOINE, 1878) described by Matthes (1977) from the « untere Unterkohle ›› turns out te be in fact a Phenacodont. Thus the decisive argument for classifying the « untere Unterkohle ›› as Lower Eocene has to be dropped. Biostratigraphically the « Unterkohle ›› and the «Basishauptrnittel ›› correspond with the lower Middle Eocene (mammal level of Messel), whereas the «unteres Hauptmittel ›› and the « untere Mittelkohle ›› are equivalent to the middle part of the middle Eocene (mammal level of lssel), and the « obere Mittelkohle ›› together with the « oberes Hauptmittel ›› coincide with the upper Middle Eocene (mammal level of Bouxwiller). 

A recently discovered, almost complete specimen of a hybodont tooth, allows us to describe as a new species, fairly common, but usually fragmentary teeth in the Navesink Formation of New Jersey: Hybodus novojerseyensis nov. sp.
Three additional taxa are also described : Heterodontus creamridgensis, Squalicorax sp. and Pseudocorax affinis which is noted for the first time in North America.
The whole elasmobranch fauna of the Early Maastrichtian localities of New Jersey is reevaluated and several generic assignments are changed in comparison to the list previously published in 1975 (Cappetta
& Case). 

About the early radiation of Rodents, we have now from the early tertiary of Asia, a new fossil record, and we can do new interpretations. First the problem of the origin of Rodents is studied : considered as a sister group of Lagomorpha, it is possible to find their ancestors between the Mixodontia. Second the new facts about the origin of modern groups of Rodents are reviewed. A general scheme of this diversification can be proposed. 

Fourteen distinct phyletical lineages which belong at least in three families: Ischyromyidae ALSTON, 1876, Gliridae THOMAS, 1896 and Theridomyidae ALSTON, 1876, have been identified after the study of more than 3600 rodent dental remains from about twenty Early and Middle Eocene european localities. A systematical and phylogenetical revision of these rodents has been achieved. Nearly all the specific and generic diagnosis are emended. Several new combinations and synonymies are proposed. Four new species and two new genera, Euromys nov. (Ailuravinae) and Hartenbergeromys nov. (Microparamyini), are named and described. Euromys nov. gen. is known by three distinctive ypresian (MP 7 to MP 10 european reference levels) chronospecies. This new lineage is thought to be the direct ancestor of Meldimys MICHAUX, 1968 and Ailuravus RUTIMEYER, 1891. A new species of the genus Plesiarctomys BRAVARD, 1850, Pl. lapicidinarum from Condé-en-Brie (MP 8-9 reference level), allows to relate the Plesiarctomys lineage to the Pseudoparamys MICHAUX, 1964 one. The taxa Sparnacomys HARTENBERGER, 1971, Pantrogna HARTENBERGER, 1971, and Corbarimys MARANDAT, 1989 are erected to genus rank; the last one is not thought to be an Ischyromyidae. A new chronospecies of Pantrogna, P. marandati nov. sp. from the locality of Prémontré (MP 10 reference level), is described. This lineage is at the origin of two others, namely Masillamys TOBIEN, 1954, including M. mattaueri (HARTENBERGER, 1975) nov. comb. (MP 10 reference level), and Hartenbergeromys nov. gen., known from MP 10 (H. hautefeuillei nov. sp.) and MP 11 (H. parvus TOBIEN, 1954) reference levels. The phylogenetical position of Hartenbergeromys nov. gen., at the origin of the european family Theridomyidae, is discussed. The systematical and phylogenetical status of two probable Paramyinae, "Paramys" woodi MICHAUX, 1964 and an unnamed genus and species, are discussed. New populations of the primitive Gliridae Eogliravus HARTENBERGER, 1971 and of the primitive Theridomyidae Protadelomys HARTENBERGER, 1968, are described and assigned to previously known species.

  

A new European rodent from the middle Eocene of Spain, Zamoramys extraneus n. gen., n. sp., appears to be closely related to the middle Eocene chapattimyid rodents of Indo-Pakistan. This contradicts the generally accepted paleobiogeographic hypothesis of a Tethyian barrier between Europe and Asia isolating Europe during the middle Eocene. Because of this barrier, some authors have proposed that European and Asian rodents were not closely related, their similarity being the result of morphological convergence. Here monophyly has been tested, using the parsimony criterion, based on an analysis of dental characters (including discussing of homology and the validity of some characteristics). Our results indicate a phylogenetic relationship among the Asiatic Ctenodactyloidea, Zamoramys from Spain, and the European endemic Theridomyoidea. We also conclude from our analysis that theridomyoids and European ischyromyoids are probably not closely related phylogenetically.