A skull of Parabos cordieri, undamaged by fossilisation, allows us to make an endocranial cast, which is compared to those of Bos taurus, Boselaphus tragocamelus and Hippotragus equinus. These are studied by general form (shape, volume, « rolling-up›› and inclination), and by detailed morphology (gyri and sulci patterns, rete mirabile, importance of rhinencephalon and cerebellum).
A preliminary study is made in order to recognize convolution topography, gyri, sulci, and the various encephalic regions as clearly on an endocranial cast as on a bare encephalon; an intermediate stage is the study of an encephalon covered with «soft» meninges.
The comparative study reveals great similarity between the endocranial cast of Parabos cordieri and that of Boselaphus tragocamelus, both belonging to the tribe Boselaphini. On the other hand, a marked differences separates the cast of Parabos cordieri from that of Bos taurus, representing the tribe Bovini to which Parabos cordieri had been referred. The endocranial cast of Hippotragus equinus, which had long been considered the descendant of Parabos cordieri, is also quite different from that of Parabos. 

The re-examination of six fossil selachian species has involved the creation of six new genera: Squatiscyllium, Protolamna, Parotodus, Abdounia, Physogaleus, Hypolophodon and of one new species : Prozolamna sokolovi. The modification of the generic statute of these species allows to clarify their systematic position and to define their relationships at a familial level. 

Nyctereutes megamastoides (Pomel) from the Villafranchian of the Auvergne and from Saint-Vallier presents cranial and dental characters sufficiently close to those of the late Pliocene canid from Perpignan (Roussillon), described by Depéret under the specific name of Canis donnezani belonging to the same genus Nyctereutes. The extinction of the European "Nyctereutes" group seems due to the too great alimentary specialization of this canid, whereas the Asiatic lineage represented in the Villafranchian by Nyctereutes sinensis Schlosser and at present by Nyctereutes procyonider Gray was able to maintain itself probably by means of a profound change in its alimentary regime. 

Cet article explore les pistes fossiles de vertébrés du Stormberg (Trias, Afrique du Sud), révélant une diversité ichnologique inédite avec 64 ichnotypes classés en tridactyles, tétradactyles et pentadactyles. Les découvertes incluent des reptiles bipèdes comme Qemetrisauropus et Prototrisauropus, dont les empreintes suggèrent des adaptations évolutives vers la bipédie, ainsi que des pentadactyles géants comme Pentasauropus, annonçant les futurs sauropodes. L’étude combine ichnologie et restes osseux pour reconstituer des écosystèmes triasiques riches, où flore (fougères, conifères) et faune témoignent d’une transition écologique majeure avant l’essor des dinosaures. 

Designated the type-locality of a late Eocene paleomammal zone, Robiac has recently been the object of important excavations. The first results of the new collecting, as  well as a revision of the material in old collections, are given in this work.
Two stratigraphic section, cutting through the two sites presently distinguished (Robiac-Nord and Robiac-Sud) reveal the lithologic variation at the base of the Fons Limestone and the localization of three fossiliferous beds at Robiac-Sud.
The molluscan fauna and the flora (charophytes), which have already been described, as well as that of the lower vertebrates, have been listed.
A list of 46 mammalian species (only 16 species were known previous to 1964) has been established. The micro-mammals, nearly all new in this fauna (marsupials, insectivores, bats, rodents, primates, and some smallsized artiodactyls), Were obtained only after screen-washing of the matrix; about 4 tons of sediment were thus treated.
The artiodactyls have been the most extensively analyzed; 6 genera, of which one is new, have been recognized. The latter is described as Robiacina minum n.g., n.sp., and represents a very small artiodactyl of the family Anoplotheriidae. The taxonomie status of certain species formerly described has been clarified by the designation of lectotypes (Cebochoerus robiucensis, Catodonerium robiacense, Xiphodon castrense).
The paleontologic corrélations at present possible between the late Eocene faunas have allowed the relative positions of te different French localities of this age to be established; the Guépelle locality, it seems, could define in the future a new paleomammal zone.
  

The eggshell of Nicobar scrubfowl (Megapodius nicobariensis) is described for the first time. Its egg porosity is calculated and discussed with data from several taxa (another megapode, some extant and fossil reptiles including a titanosaur group) in order to compare incubation types with eggshell structure. Eggshell microstructure reflects first phylogenetic traits and does not seem to have developed major adaptative features due to the incubation conditions, except for the pore canals. 

The presence of cambaytheres, the sister group of perissodactyls, in western India near or before the time of collision with Asia suggests that Perissodactyla may have originated on the Indian Plate during its final drift towards Asia. Herein we reinforce this hypothesis by reporting two teeth of the first early Eocene tapiromorph Perissodactyla from the Cambay Shale Formation of Vastan Lignite Mine (c. 54.5 Ma), Gujarat, western India, which we allocate to a new genus and species, Vastanolophus holbrooki. It presents plesiomorphic characters typical of the paraphyletic “Isectolophidae,” such as small size and weak lophodonty. However, the weaker hypoconulid and low paralophid, higher cusps, lower cristid obliqua, and the lingual opening of the talonid are found in Helaletidae, the most primitive tapiroid family. V. holbrooki, gen. et sp. nov., may be the oldest and the most primitive tapiroid, suggesting that at least tapiroid perissodactyls originated on India. 

The fauna studied in the following work involves the dentitions and skulls more or less complete of 5 genera, among which only Cephalomys was previously known by its skull. One must notice that the Salla's species of this genus is a new one. Sallamys, rather small, shows a dentition rather similar to that of Platypittamys Wood from Patagonia. The upper molars, more primitive than those of this last genus, according to the smaller dimensions of the hypocone, retain a distinct metaloph. This metaloph tends to be reduced in a way which may give us a possibility to understand how it disappeared in Platypittamys. The upper P4 can be compared as well to that of Platypittamys as to that of Gaudeamus from the African Oligocene. The lower P4, more molarized than that of Platypittamys, is already moving towards the miocene type of structure. The infraorbital foramen is wide and the insertion of the masseter on the muzzle is spacious. Branisamys, genus of a great size, shows an auditory region partly preserved, peculiarly the promontorium with the fenestra rotunda, entirely of the Hystricognathi type. Upper molars are very clearly pentalophodont. A new reconstruction is proposed for the tooth called Villarroelomys by Hartenberger. This tooth is shown to be a lower D4, perhaps of Branisamys , certainly of a rather nearly allied form, and Hartenberger does agree with the essential part of this new conclusion. Of Incamys, two incomplete skulls are known, each one being admitted to be the type of a distinct species, the first one being I. bolivianus, I. pretiosus the second. The infraorbital foramen is of a great size and the impression of the masseter on the muzzle is spacious. The sphenopalatine foramen is widely developed and of a really very uncommon great size. Only Thryonomys from Africa shows a similar tendency to the enlargement of this foramen, but not so extreme. The main basicranial foramina can be observed. The upper teeth, hemi-hypsodont, show, either a vestigial metaloph, similar to that of recent Thryonomys from Africa, associated with a well developed mesoloph, either a well developed metaloph, while the mesoloph is reduced or absent. Cephalomys was previously known by anterior parts of the skull showing a wide infraorbital foramen and a spacious facial insertion of the masseter. Its lacrymal is of the phiomorph type and the spheno-palatine foramen is seemingly of great size, like in Incamys. The species is new. The varied peculiarities of the upper teeth of these genera can be easily understood if we refer to the plan of the teeth of Phiomys andrewsi from the Oligocene and Miocene of Africa. The structure of this genus, clearly more primitive, still typically brachyodont, shows and clearly explains the fundamental coherence of the varied realisations arised from such a structure. Luribayomys n.g. is known only by an anterior half of a skull without teeth. It is remarquable by the great development of the masseter's insertions on the muzzle and by the lacrymal region, well preserved, typically phiomorphid. The classification previously published by A.E. Wood and B. Patterson is granted in its essential parts, provisionally, but not as a definitive solution. Nevertheless the Dasyproctidae are integrated within the Cavioidea, following the conclusions of Bugge and of Vucetich, reached independently. The conclusion emphasizes the exceptional meaning of the fauna of Salla-Luribay. This shows that Platypittamys, while interesting, can no more be supposed certainly representative of the normal structure of the Oligocene Caviomorph, and not even of their ancestors. The anatomical peculiarities exhibited in these new samples, auditory region, lacrymal, spheno-palatine foramen, reinforce the primitive structural identity with the Phiomorpha. Similarly, the new lower D4 favour very close relationships, ever if the affinities of the D4 has been questioned or minimized by Wood and Patterson. It is certainly possible to admit that parallelism could explain limited similarities, like the presence in North America of Rodents with an hystricomorph type of infraorbital foramen and an hystricognath mandible. But if the parallelism could be a sufficient explanation of the identical association of multiple and complete structures observed in the Caviomorpha and Phiomorpha, all the Zoological systematic would have to be questioned. The last positions of A.E. Wood on the subject (1975) are revised and criticised, and the recent publications studying the problems of distance between Africa and South America in Eocene time, as a consequence of the drift, are quoted; the possibility of transportation by rafts is shown. A new hypothesis is proposed about the interrelationships of Pentalophodont Rodents, with interesting paleobiogeographic implications. 

Abundant crocodilian fossil remains have been collected, mainly by R. Lavocat. from several levels of the
Palaeogene of the Tilemsi valley (eastern Mali). Several dyrosaurids occur in the Palaeocene (Montian ?) : Phosphatosaurus sp., Hyposaurus bequaerti (Dollo). H. wilsoni (Swinton). H. nopesai (Swinton), Rhabdognathus rarus Swinton, R. compressus n. sp. ln the lower Eocene has been found a dyrosaurid which cannot be accurutely determined.In the middle Eocene were found a dyrosaurid, Tilemsisuchus lavocati n. g., n. sp., and an indeterminate eusuchian. 

New and hitherto unpublished mammals from the stratigraphical levels Unterkohle, Untere Mittelkohle and Obere Mittelkohle of the Geiseltal near Halle, GDR, are described (= biochronological levels MP 11-13, Geiseltalian sensu Franzen & Haubold 1986a, b). The marsupial taxa Amphiperatherium aff. maximum (MP 12), A. goethei (MP 12), and Peratherium aff. monspeliense (MP 12 and 13) are recorded for the first time. A lectotype for Amphiperatherium giselense is designated, and the alleged primate Microtarsioides voigzi is assigned to Marsupialia, incertae sedis. A new insectivore species, Saturninia ceciliensis n. sp., is described (MP 13). The anteater Eurotamandua joresi is recorded for the first time outside its type locality, Grube Messel, FRG (MP 11). The present humerus and ulna display the autapomorphic features of the myrmecophagids. 

New faunal and stratigraphical data on the vertebrates localities from the early Paleogene of the Ouarzazate Basin (Adrar Mgorn 1, Adrar Mgorn 1 bis et N'Tagourt 2), Morocco, are presented. A magnetostratigraphical study, the first for such early Paleogene Arabo-African mammal localities, and the discovery of probable remains of the nannofossil Discoaster support the Thanetian age of the Adrar Mgorn 1 site. The magnetostratigraphy suggests a slightly later age than was thought for the Paleogene formations of the local series of Tinerhir and for the vertebrate localities: late or latest Thanetian for Adrar Mgorn 1 and Adrar Mgorn 1 bis, middle Ypresian for N'Tagourt 2. It also indicates a lower position of the KT boundary in the series. Two tons of matrix recovered in the vertebrate sites have vielded new data on the micromammals. A damaged lower molar from N'Tagourt 2 is referable to Khamsaconus bulbosus  and supports the proboscidean affinities of this species and especially possible relationships with bunolophodont taxa such as elephantiforms. A lower molar from Adrar Mgorn 1 bis belongs to a new form which can be identified as a plesiadapiform or an euprimate close to Altiatlasius koulchii though significantly larger. A new material from Adrar Mgorn 1 illustrates a new dilambdodont adapisoriculid species which is referable to Garatherium : ?Garatherium todrae n. sp. Another species referred to Garatherium is known in the locality (?Garatherium n. sp.). Garatherium is a new lineage from the Ouarzazate basin which crosses the Paleocene-Eocene boundary together with Palaeoryctes, Didelphodontinae gen. and sp. 2, Todralestes, and Afrodon, and it is the first Paleocene-Eocene lineage identified outside of this basin (Garatheríum is based on a species from El Kohol, Algeria). Among the Paleocene-Eocene lineages from the Ouarzazate basin, it should be also mentioned a new possible carnassial form (carnivoran or creodont; Adrar Mgorn 1), and an upper molar of Cimolestes cf. incisus (Adrar Mgorn 1 bis). The upper molar THR 168 previously reported as from an indeterminate didelphodontine is here identified as the M1/ of Afrodon chleuhi. The micromammal faunas from the Ouarzazate basin are positioned in the global chronological framework of the mammal localities from the Paleogene of the Arabo-African domain. 

The earliest North American rodents occur in basal Clarkforkian beds of the Fort Union Formation at Big Multi Quarry near Bitter Creek, northern Washakie Basin, Sweetwater County, Wyoming, and in closely correlative Fort Union beds formerly accessible in the Eagle Coal Mine near Bear Creek, northern Clark's Fork Basin, Carbon County, Montana. Two new species of early Clarkforkian rodents, Paramys adamus and Alagomys russelli, are described from Big Multi Quarry. Paramys adamus is represented by virtually complete upper and lower dentitions, which demonstrate that this species is one of the most primitive North American paramyids yet discovered. These specimens form the basis for a reevaluation of the content and stratigraphic range of P. atavus, which is known with certainty only from Bear Creek. Alagomys russelli is the first North American record for the enigmatic rodent family Alagomyidae, otherwise known from ?late Paleocene-early Eocene localities in Mongolia and China. Phylogenetic analysis of dental and gnathic traits suggests that Alagomyidae form the sister group of all other undoubted rodents. At least two rodent clades, alagomyids and basal paramyids, seem to have invaded North America from Asia at the beginning of Clarkforkian time, but only the paramyids persisted to undergo a significant evolutionary radiation in North America. 

A new genus and species of haplobunodontid artiodactyl, Hallebune krumbiegeli nov. gen. nov. sp., is described from the middle Eocene Geiseltal Fauna (Saxon-Anhalt, Germany). Its stratigraphic range is restricted to the MP 13 ("obere Mittelkohle") of the Geiseltalian. The material consists of seven fragments, representing both upper and lower jaws. It is indicated, that H. krumbiegeli was a small bunodont herbivore. The size and primitive character of the teeth suggest that the genus possibly is directly derived from Diacodexeidae.The new material forms the basis for reinterpreting the haplobunodontid phylogeny. 

Voici une centaine d'années débutait sur les Causses du Quercy l'exploitation des phosphorites qui devait permettre d'amasser des collections de Vertébrés fossiles uniques au monde. Ces fossiles firent l'objet de nombreuses publications et les noms de Filhol, Gaudry, Schlosser, Stehlin, Teilhard de Chardin sont attachés à ces premières études.

Après quelques années de prospection, les fouilles dans le Quercy ont repris en 1965, organisées par les laboratoires montpelliérains et parisiens. Ces premières recherches ont été aidées par le Centre National de la Recherche Scientifique qui a organisé une « table ronde» pour faire le point des premiers résultats obtenus. Ceux-ci ont été jugés suffisamment prometteurs pour que le C.N.R.S, ait décidé de poursuivre son effort par l'intermédiaire d'une Recherche Coopérative sur Programme.



  

Functional morphology invertebrate paleontology, édité par Jeffrey J. THOMASON, 1995. Cambridge University Press, xi + 277 p. ISBN 0-521-44095-5, f. 45,00 ($ 69,95).

La morphologie fonctionnelle appliquée aux fossiles demeure un sujet à la fois passionnant et important. Passionnant, car elle s'adresse à des êtres disparus, aux combinaisons de caractères parfois inattendues et sans contrepartie chez les formes actuelles. Sujet important, car il faut, au delà de la constatation classique de l'adéquation de la forme des organes à leur fonction, expliquer l'origine des structures par évolution. L'évolution ou descendance avec modification, implique que les êtres vivants sont reliés les uns aux autres et qu'i! est possible de tenter à l'aide d'un raisonnement par inférence, la caractérisation des ancêtres en fonction des traits des formes auxquelles ils ont donné naissance. Mais il faut ajouter que par sa fonction, la structure relie immédiatement l'animal à son milieu puisqu'elle assure un rôle biologique. 

The genus Parasteiromys AMEGHINO, 1904 is revalidated, and P. friantae sp. nov. (Hystricognathi, Erethizontidae) from Colhuehuapian (early Miocene) sediments of the southern cliff of Colhue-Huapi Lake (Province of Chubut, Argentina), is described. The molar morphology of these taxa and of living porcupines adds new elements to understand the dental evolution of the Erethizontidae, and to propose the hypothetical ancestral molar pattern for this family. This pattern does not correspond to any of the morphologies traditionally proposed as ancestral for South American hystricognathous rodents. The proposed pattern is characterized by a metaloph disconnected from the posteroloph and oriented towards the hypocone, and the third loph incompletely developed with the lingual portion homologous to the mesolophule of Baluchimyinae (Chapattimyidae) from the Miocene of Pakistan. The inferred steps of the molar evolution of erethizontids towards the pentalophodont condition, considered derived for the family, are illustrated. This study strengthens the hypothesis placing erethizontids in a basal position among rodents of the suborder Hystricognathi.

  

The revision of the old collections of fossil birds from the “Phosphorites du Quercy” and the study of new material give the following results (Gruiformes, Cariamae) :  The humeri and most of the carpometacarpi described under the name Filholornis belong in Elaphrocnemus. The ulnae ascribed to Fïlholornis belong in Idiornis. Most of the post-cranial elements of the genera Elaphrocnemus and Idiornis are described and show great similarities with recent Cariamidae and Opisthocomidae, and fossil Bathornithinae.  A new genus and a new species, Oblitavis insolitus, are created in the sub-family Idiornithinae; two new species are described in the genera Elaphrocnemus (E. brodkorbz) and Idiornis (I. itardiensis), and the species Elaphrocnemus gracilis is transferred to the genus Idiornis.  The genus Propelargus Lydekker is transferred from the family Ciconiidae to Cariamidae.  A new generic name, Occitaniavis, is created for the species Geranopsis elatus, which belong in Cariamidae, while the type-species of the genus, Geranopsis hastingsiae, is a member of the Gruidae.  The affinities between the Quercy avifauna and the Neotropical one is emphasized by the occurrence of Phorusrhacidae, previously known only from the Cenozoic of South America and the Late Pliocene or Early Pleistocene of North America. Thanks to the material collected during the new excavations, the stratigraphical position of most of the species is stated precisely, and evolutionary lineages are outlined. This study shows that the suborder Cariamae, presently restricted to two South American genera, was already extremely diversified during the Eocene, and widespread in Europe and North America. 

The comparative morphology and biometry of the lower cheek teeth of modern Equus are studied on approximately 300 mandibles belonging to the 10 usually recognised species : Equus grevyi, E. burchelli, E. quagga, E. zebra, E. africanus, E. asinus, E. hemionus, E. klang, E. przewalskii, E. caballus. The studied parameters comprise : occlusal length and width, postflexid length and index ; shape of the double knot (metaconid + metastylid + lingual groove) ; depth of the vestibular groove on the molars ; frequency of the pli caballinid, protostylid and other enamel plications or islets ; frequency of the dP/l.
The same methods of study are applied to a number of North American, Eurasian and African species. For the sake of comparison, some Hemphillian equids were observed (Dinohippus interpolatus, Dinohippus leidyanus, Astrohippus ansae, Phiohippus mexicanus) but most of the discussed material belongs to Pliocene or Pleistocene species of Equus : the « stenonine ›› E. stenonis, E. simplicidens, E. sanmeniensis and E. teilhardi; the « caballine ›› E. scotti, E. lambei, E. Iaurentius, E. mosbachensis, E. germanicus, E. gallicus, E. taubachensis and the Liakhov horse. The relationships of other species, in particular the North American E. calobatus, E. occidentalis, E. cf mexicanus are not clear for the moment. ln Africa, the Plio-Pleistocene species from Koobi Fora (Kenya) show some stenonine and perhaps asinine affinities. The relationships of E. numidicus and E. tabeti are uncertain but these species are probably related to the East African ones. E. mauritanicus is most certainly related to the Quagga group.
The biometrical data are gathered in 32 tables ; 4 photographie plates and 19 figures illustrate the next. The whole is a complement of the previously published studies of the skulls, upper cheek teeth, incisors and metapodials of modern and fossil Equus.

  

The Montian locality of Hainin (Hainaut, Belgium) yielded about twenty teeth of Multituberculates. They are very peculiar forms, showing no affinities, at the generic level, with those hitherto known from North America, Asia and Europe. They are referred to the new taxa Boffius splendidus nov. gen., nov. sp., Hainina belgica nov. gen., nov. sp., and H. godfriauxi nov. gen., nov. sp. They expose some common features, such as the advanced type of first upper molar. possessing at least three complete rows of cusps. Because of this, and also of the upper premolar reduction, Boffius splendidus appears as the most specialized form within the Ptilodontoidea suborder.
Several other characters of Hainina seem to be less advanced, such as the great number of upper premolars and the simple cusp-formula of the first lower molar.
Till now, only H. godfriauxi has been recovered within the Thanetian fauna from Cemay-les-Reims, where it is very poorly documented. 

Les Pistes de Vertébrés du Stormberg Supérieur ("Trias terminal à Rhétien"), ou Quthingien

Si les zones du Stormberg inférieur se sont révélées contenir de nombreuses traces, surtout dans les faciès dits "Molteno moyen et supérieur", représentant apparemment la base du Keuper, il est frappant de voir pratiquement l'ensemble de cette grosse faune "Molteno" disparaître avec la fin de cette période, que nous avons appelée le "Maphutsengien".

Dès les premières zones du Stormberg supérieur, que nous nommons le"Quthingien" la zoocénose et la phytocénose, en même temps que les données d'ensemble manifestées par l'environnement, sont modifiées. Nous ne verrons plus guère de dépôts marécageux à flore riche et variée, parfois même luxuriante. Les fougères elles-mêmes ont disparu. Elles sont remplacées par de maigres plantes, aux feuilles très souvent filiformes qui paraissent témoigner d'un climat continental. Le sol est devenu de plus en plus rouge, avec des variations latérales beaucoup plus accusées. Les fleuves amenant des galets des monts du "Grand Sud" ont tari. La faune va en subir les conséquences. Certaines des espèces se révèleront sautillantes ou coureuses, pour un grand nombre plus légères et pour la quasi-totalité d'apparence carnivore ou entomophages, les phytophages devant se contenter d'un régime ingrat,difficile ou à tout le moins irrégulier,les dépôts le montrent.

C'est dans ces conditions que s'inaugure notre Etage nouveau,quelque peu discordant sur les zones A/5, A/6 ou A/7 du Stormberg inférieur (Maphutsengien). Le Stormberg supérieur (ou Quthingien) commence avec le paléopaysage remarquable dit de Moyeni, que nous allons maintenant étudier, typologiquement, avec ses homologues du même âge. Quelques 38 types d'animaux tous nouveaux vont défiler à nos yeux lors de la zone de base de cet Etage, ou zone B/1.

L'on nous avait proposé d'intituler ce Ile Tome de la série : "La grande Dalle de Moyeni et ses homologues. Paléo-spectacles, scènes et paysages animaux au Lesotho à l'approche du Trias finissant". Nous avons préféré garder le sous-titre plus haut, peut-être plus prosaïque.

Un llle Tome est en préparation : "Les développements ultérieurs et terminaux de la faune du Gondwana".